Rainey, Paul B., and Benjamin Kerr. “Conflicts among levels of selection as fuel for the evolution of individuality.” The major transitions in evolution revisited (2011): 141-162.
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This chapter describes the evolutionary transition from single cells to multicellular individuals. It shows that in multicellular organisms, individuality is ultimately a property of individual cells that give rise to a multicellular organism in each generation. It then investigates the breakdown of cooperation, in the course of developing a heterodox idea on the origins of the germ-soma distinction. It examines the evolution of multicellular development in the Pseudomonas system by introducing a discrete-time theoretical model. The described unconventional life cycles that span the multilevel selection (MLS)-1 to MLS-2 juncture are founded in experimental reality.